From the tiny acorn grows the mighty oak and the acorn stem is the link that holds the two together until the fall ripening. The acorn cap with stem intact provides budding tree taxonomists with important clues to the true species identity of its source within the oak family.

View a low magnification image of the acorn stem.

View a medium magnification image of the acorn stem.

Classified as a nut, because of its bony pericarp, the acorn is actually the fruit of the oak tree. Particular to the genus Quercus, the stem broadens to the cupule or cap that holds the oak seed and fruit in place. The burr oak, Q. macrocarpa, so named because of its characteristic large seeds or acorns, was known as u'tahu can in the Native American tongue of the Lakota people, meaning acorn stem tree. More than half of the one-inch acorn is enclosed in a fringed cupule. Valued as an important diet item with great storage capability and mobility, Native Americans made acorns much more palatable by first blanching the oak nuts with wood ashes to remove a bitter taste associated with tannins.

Wildlife, including turkeys, blue jays, squirrels, and deer, utilize acorns as a food source that is rich in carbohydrates and fats. Cached acorns for future feedings are often forgotten and may germinate into new oak trees. Acorns and oak trees have long been part of superstition and religions, such as the Druids of the ancient British Isles. It was believed that an acorn should be carried to bring luck to the bearer and ensure a longer life. Others believed that placing an acorn at the window would ward off lightning.

Some acorns, such as those of the swamp white oak (Q. bicolor), found in the Ozarks and northern Missouri, are usually carried in pairs, characterized by a relatively long stem that is 2.5 inches long, and a scaly cup that is covered with fine, wooly "hair". Other varieties, such as the fruit of the black oak (Q. velutina), require two growing seasons to ripen, and then bear kernels that are extremely bitter. No matter what reproductive strategy has evolved for a particular oak species, the stem of the acorn must balance the task of supporting the heavy nut, coupled to a breaking tolerance that facilitates the success of nut gatherers and unsuspecting oak tree dispersers (such as squirrels and blue jays).

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